Laboratory of Integrative and Comparative Herpetology
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LICH Publications

  1. Espinoza, R. E. and F. Lobo. 1996. Possible communal nesting in two species of Liolaemus lizards (Iguania: Tropiduridae) from northern Argentina. Herpetological Natural History 4:65–68.
  2. Etheridge, R. and R. E. Espinoza. 1997. New records and natural history notes for lizards from northwestern Argentina. Herpetological Review 28:160–161.
  3. Espinoza, R. E. and C. R. Tracy. 1997. Thermal biology, metabolism, and hibernation. Pp. 149–184, in: The Biology, Husbandry and Health Care of Reptiles. L. J. Ackerman (Ed.). Volume 1, Biology of Reptiles. TFH Publications, Neptune City, New Jersey.
  4. Espinoza, R. E., C. R. Tracy, and C. R. Tracy. 1998. A safe, single-application procedure for eradicating mites on reptiles. Herpetological Review 29:35–36.
  5. Espinoza, R. E., C. A. Carreño, C. R. Tracy, and C. R. Tracy. 1998. Geographic distribution. Sauromalus obesus obesus. Herpetological Review 29:51–52.
  6. Nussear, K. E., R. E. Espinoza, C. M. Gubbins, K. J. Field, and J. P. Hayes. 1998. Diet quality does not affect resting metabolic rate or body temperatures selected by an herbivorous lizard. Journal of Comparative Physiology B 168:183–189.
  7. Lobo, F. and R. E. Espinoza. 1999. Two new cryptic species of Liolaemus (Iguania: Tropiduridae) from northwestern Argentina: resolution of the purported reproductive bimodality of Liolaemus alticolor. Copeia 1999:122–140.
  8. Espinoza, R. E., C. A. Carreño, and R. W. Van Heest. 1999. Geographic distribution. Elgaria coerulea. Herpetological Review 30:51–52.
  9. Espinoza, R. E. 1999. Reptiles and herbivory. Herpetological Review 30:125–126.
  10. Halloy, M. and R. E. Espinoza. 1999–2000. Territorial encounters and threat displays in the Neotropical frog, Phyllomedusa sauvagii (Anura: Hylidae). Herpetological Natural History 7:175–180.
  11. Schulte, J. A. II, J. R. Macey, R. E. Espinoza, and A. Larson. 2000. Phylogenetic relationships in the iguanid lizard genus Liolaemus: multiple origins of viviparous reproduction and evidence for recurring Andean vicariance and dispersal. Biological Journal of the Linnean Society 69:75–102.
  12. Espinoza, R. E., F. Lobo, and F. B. Cruz. 2000. Liolaemus heliodermis, a new lizard from northwestern Argentina with remarks on the content of the elongatus group (Iguania: Tropiduridae). Herpetologica 56:235–244.
  13. Simandle, E. T., R. E. Espinoza, K. E. Nussear, and C. R. Tracy. 2001. Lizards, lipids, and dietary links to animal function. Physiological and Biochemical Zoology 74:625–640.
  14. Etheridge, R. and R. E. Espinoza. 2000. Taxonomy of the Liolaeminae (Squamata: Iguania: Tropiduridae) and a semi-annotated bibliography. Smithsonian Herpetological Information Service 126:1–64.
  15. Scheltinga, D. M., B. G. M. Jamieson, R. E. Espinoza, and K. S. Orrell. 2001. Descriptions of the mature spermatozoa of the lizards Crotaphytus bicinctores, Gambelia wislizenii (Crotaphytidae), and Anolis carolinensis (Polychrotidae) (Reptilia, Squamata, Iguania). Journal of Morphology 247:160–171.
  16. Cooper, W. E. Jr., J. J. Habegger, and R. E. Espinoza. 2001. Responses to prey and plant chemicals by three iguanian lizards: relationship to plants in the diet. Amphibia-Reptilia 22:349–362.
  17. Espinoza, R. E. and F. B. Cruz. 2003. Hyla andina. Diet. Herpetological Review 34:136.
  18. Espinoza, R. E. 2003. Comment on the proposed conservation of usage of the names Phymaturus Gravenhorst, 1837 and Lacerta palluma Molina, 1782 (currently Phymaturus palluma; Reptilia, Sauria) by designation of a neotype for Lacerta palluma Molina, 1782. Bulletin of Zoological Nomenclature 60:58.
  19. Espinoza, R. E. and F. Lobo. 2003. Two new species of Liolaemus lizards from northwestern Argentina: speciation within the northern subclade of the elongatus group (Iguania: Liolaemidae). Herpetologica 59:89–105.
  20. Hernandez, R. A. and R. E. Espinoza. 2004. Description of the female of the enigmatic lizard, Liolaemus heliodermis (Iguania: Liolaemidae). Herpetological Review 35:227–229.
  21. Espinoza, R. E., J. J. Wiens, and C. R. Tracy. 2004. Recurrent evolution of herbivory in small, cold-climate lizards: breaking the ecophysiological rules of reptilian herbivory. Proceedings of the National Academy of Sciences, USA 101:16819–16824.
  22. Lobo, F. and R. E. Espinoza. 2004. Two new Liolaemus from the Puna region of Argentina and Chile: further resolution of purported reproductive bimodality in Liolaemus alticolor (Iguania: Liolaemidae). Copeia 2004:850–866.
  23. Cruz, F. B., L. A. Fitzgerald, R. E. Espinoza, and Schulte, J. A. II. 2005. The importance of phylogenetic scale in tests of Bergmann’s and Rapoport’s rules: lessons from a clade of South American lizards. Journal of Evolutionary Biology 18:1559–1574.
  24. Tracy, C. R., K. M. Flack, L. C. Zimmerman, R. E. Espinoza, and C. R. Tracy. 2005. Herbivory imposes constraints on voluntary hypothermia in lizards. Copeia 2005:12–19.
  25. Tracy, C. R., K. E. Nussear, T. C. Esque, K. Dean-Bradley, C. R. Tracy, L. A. DeFalco, K. T. Castle, L. C. Zimmerman, R. E. Espinoza, and A. M. Barber. 2006. The importance of physiological ecology in conservation biology. Integrative and Comparative Biology 46:1191–1205.
  26. Lancaster, J. R., P. Wilson, and R. E. Espinoza. 2006. Physiological benefits as precursors of sociality: why banded geckos band. Animal Behaviour 72:199–207.
  27. Espinoza, R. E. and K. de Queiroz. 2008. Historical perspectives: Richard Emmett Etheridge. Copeia 2008:708–717.
  28. Abdala, C. S., A. S. Quinteros, and R. E. Espinoza. 2008. Two new species of Liolaemus (Iguania: Liolaemidae) from the Puna of northwestern Argentina. Herpetologica 64:458–471.
  29. Espinoza, R. E. and S. Quinteros. 2008. A hot knot of toads: aggregation provides thermal benefits to metamorphic Andean toads. Journal of Thermal Biology 33:67–75.
  30. Espinoza, R. E. 2009. Western Fence Lizard (Sceloporus occidentalis). Pp. 234–237, in: Lizards of the American Southwest: A Photographic Fieldguide, L. L. C. Jones and R. E. Lovich (eds.). Rio Nuevo Press, Tucson, Arizona.
  31. Espinoza, R. E. 2009. Mohave Fringe-toed Lizard (Uma scoparia). Pp. 278–281, in: Lizards of the American Southwest: A Photographic Fieldguide, L. L. C. Jones and R. E. Lovich (eds.). Rio Nuevo Press, Tucson, Arizona.
  32. Espinoza, R. E. 2009. Western Banded Gecko (Coleonyx variegatus). Pp. 312–315, in: Lizards of the American Southwest: A Photographic Fieldguide. L. L. C. Jones and R. E. Lovich (eds.). Rio Nuevo Press, Tucson, Arizona.
  33. Lobo, F., R. E. Espinoza, and A. S. Quinteros. 2010. A critical review and systematic discussion of recent classification proposals for liolaemid lizards. Zootaxa 2549:1–30. ​
  34. Espinoza, R. E., F. Lobo, and R. Etheridge. 2011. Taxonomic history of the iguanian lizard Liolaemus pictus major Boulenger with a revalidation of Liolaemus capillitas Hulse. Journal of Herpetology 45:129–133.
  35. Lobo, F. and R. E. Espinoza, E. A. Sanabria, and L. B. Quiroga. 2012. A new Phymaturus (Iguania: Liolaemidae) from the southern extreme of the Argentine Puna. Copeia 2012:12–22.
  36. Valdecantos, M. S., F. Arias, and R. E. Espinoza. 2012. Herbivory in Liolaemus poecilochromus, a small, cold-climate lizard from the Andes of Argentina. Copeia 2012:203–210.
  37. Weeks, D. M. and R. E. Espinoza. 2013. Lizards on ice: comparative thermal tolerances of the world’s southernmost gecko. Journal of Thermal Biology 38:225–232.
  38. Böhm, M., B. Collen, J. E. M. Baillie, P. Bowles, J. Chanson, N. Cox, G. Hammerson, M. Hoffmann, S. R. Livingstone, M. Ram, A. G. J. Rhodin, S. N. Stuart, P. P. van Dijk, B. Young, + 225 authors listed alphabetically including R. E. Espinoza. 2013. The conservation status of the world’s reptiles. Biological Conservation 157:372–385.
  39. Larson, G. C., T. Wolfmeyer, B. Singh, R. E. Espinoza, C. Huntley, and G. B. Pauly. 2015. Geographic distribution. Hemidactylus turcicus. Herpetological Review 46:59.
  40. Pauly, G. B., A. Wells, M. H. Espinoza, T. A. Wake, and R. E. Espinoza. 2015. Geographic distribution. Hemidactylus turcicus. Herpetological Review 46:59.
  41. Espinoza, M. H. and R. E. Espinoza. 2015. Geographic distribution. Phelsuma laticauda. Herpetological Review 46:60.
  42. Abdala, C. S., D. Baldo, R. A. Juárez, and R. E. Espinoza. 2016. The first parthenogenetic pleurodont iguanian: a new all-female Liolaemus (Squamata: Liolaemidae) from western Argentina. Copeia 104:487–497.
  43. Altig, R., G. B. Pauly, and R. E. Espinoza. 2016. Geographic distribution. Hemidactylus turcicus. Herpetological Review 47:628.
  44. Fleischner, T. L., R. E. Espinoza, G. Gerrish, H. W. Greene, R. Wall Kimmerer, E. A. Lacey, S. Pace, J. K. Parrish, H. Swain, S. Trombulak, S. Weisberg, D. W. Winkler, and L. Zander. 2017. Teaching biology in the field: importance, challenges, and solutions. BioScience 67:558–567.
  45. Thompson, L. R., J. G. Sanders, D. McDonald, A. Amir, J. Ladau, K. J. Locey, R. J. Prill, A. Tripathi, S. M. Gibbons, G. Ackermann, J. A. Navas-Molina, S. Janssen, E. Kopylova, Y. Vázquez-Baeza, A. Gonzalez, J. T. Morton, S. Mirarab, Z. Z. Xu, L. Jiang, M. F. Haroon, J. Kanbar, Q. Zhu, S. J. Song, T. Kosciolek, N. A. Bokulich, J. Lefler, C. J. Brislawn, G. C. Humphrey, S. M. Owens, J. Hampton-Marcell, D. Berg-Lyons, V. McKenzie, N. Fierer, J. A. Fuhrman, A. Clauset, R. L. Stevens, A. Shade, K. S. Pollard, K. D. Goodwin, J. K. Jansson, J. A. Gilbert, R. Knight, and The Earth Microbiome Project Consortium [= 260 authors including R. E. Espinoza]. 2017. A communal catalogue reveals Earth’s multiscale microbial diversity. Nature 551:457–463.
  46. Grismer, L. L., P. L. Wood, Jr., M. K. Thura, E. S. H. Quah, M. S. Grismer, M. M. Murdoch, R. E. Espinoza, and A. Lin. 2018. A new Cyrtodactylus Gray, 1827 (Squamata, Gekkonidae) from the Shan Hills and the biogeography of bent-toed geckos from eastern Myanmar. Zootaxa 4446:447–500.
  47. Grismer, L. L., P. L. Wood, Jr., E. S. H. Quah, M. L. Murdoch, M. S. Grismer, M. W. Herr, R. E. Espinoza, R. M. Brown, and A. Lin. 2018. A phylogenetic taxonomy of the Cyrtodactylus peguensis (Reptilia: Squamata: Gekkonidae) with descriptions of two new species from Myanmar. PeerJ 6:e5575.
  48. Grismer, L. L., P. L. Wood, Jr., E. S. H. Quah, M. K. Thura, R. E. Espinoza, M. S. Grismer, M. L. Murdoch, and A. Lin. 2018. A new species of crocodile newt Tylototriton (Caudata: Salamandridae) from Shan State, Myanmar (Burma). Zootaxa 4500:553–573.
  49. Aragón y Traverso, J. H., E. A. Sanabria, L. B. Quiroga, and R. E. Espinoza. 2019. Siagonodon borrichianus. Predation. Herpetological Review 50:602.
  50. Grismer, L. L., P. L. Wood, Jr., E. S. H. Quah, M. K. Thura, R. E. Espinoza, M. L. Murdoch. 2019. A new species of crocodile newt Tylototriton (Caudata: Salamandridae) from northern Myanmar (Burma). Journal of Natural History 53:475–495.
  51. Stinner, J. N. and R. E. Espinoza. 2020. Crotalus oreganus helleri. Diet. Herpetological Review 51:140.
  52. Monroe, D. J., G. B. Pauly, T. C. Ruppert, and R. E. Espinoza. 2020. Historic and recent range expansion of the Mediterranean House Gecko (Hemidactylus turcicus) into the eastern USA. Herpetological Review 51:447–451.
  53. Pauly, G. B., L. Gordon, and R. E. Espinoza. 2020. Spea hammondii. Longevity. Herpetological Review 51:824–825.
  54. Weeks, D. M. and R. E. Espinoza. 2020. Warmer isn’t always better: performance eurythermy in a cold-climate gecko. Journal of Experimental Zoology 333:205–213.

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​Department of Biology
College of Science and Mathematics
California State University, Northridge

Northridge, California 91330-8303, USA
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